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On the Cover: The ultraviolet-B spectral band (UV-B, 280­315 nm) of solar radiation contributes a very small fraction of the short-wave photons received by terrestrial organisms in the natural environment. Yet plant exposure to solar UV-B can induce significant changes in growth, synthesis of secondary compounds, and plant defense against folivore insect. A special including contributions of Casati and Walbot (pp. 1739-1754), Izaguirre et al. (pp. 1755-1767), and Holley et al. (1728-1738) sheds new light into the mechanisms that mediate plant responses to solar UV-B radiation and the interactions between responses to UV-B and biotic stressors. The image shows a plant of Nicotiana longiflora (the species used in the field studies of Izaguirre et al.), growing in its native environment in central Argentina (Cover image by Carlos A. Mazza and Carlos L. Ballaré).