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On the Cover: Myrosinase or thioglucoside glucohydrolase (TGG) plays an important role in plant defense, hydrolyzing glucosinolates into products toxic to microbes and herbivores. Understanding the organization of the myrosinase-glucosinolate system provides important insights into organization of crucifer defense and signaling. TGG1 is one of the functional myrosinases in Arabidopsis. Husebye et al. (pp. 1180-1188) have mapped the spatial distribution of TGG1 in Arabidopsis. The cover shows the myrosinase-containing guard cells and phloem idioblasts in the upper flower stalk visualized by -glucuronidase enzyme staining. Located next to myrosin idioblasts of the phloem bundle are S cells, rich in glucosinolate substrate, supporting a cellular separation of myrosinase and glucosinolate substrate.