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On the Cover: We show a cross-section of a mature leaf of Flaveria bidentis immunolabeled with antibody against NADP-malic enzyme. Yellow dots indicate where the enzyme is present, demonstrating its accumulation in bundle-sheath chloroplasts of this C₄ plant, where it functions in C₄ photosynthesis. Drincovich et al. (pp. 733-744) show the occurrence and distribution of three plastid isoforms of the enzyme among C₃, C₃-C₄ intermediate, and C₄ Flaveria species, including a low-activity, 72-kD isoform that is expressed in leaf, stem, and root; a 64-kD isoform appearing in leaves of photosynthetic intermediates; and a high-activity 62-kD isoform that is dominant in leaves of C₄ species. The authors consider the evolutionary implications relative to development of C₄ photosynthesis in the genus Flaveria.